Page 204 - Proceedings of the State Natural History Museum. Issue 37 (Lviv, 2021)

Іs clytini monophyletic? The evidence from five-gene phylogenetic … 203

transverse keels or fields of coarse chitin grains. Elytra slightly attenuated posteriorly,
apically truncated, acutely spined. Legs very elongated. Femora gradually thickened
posteriorly, apically acutely spined.
Diagnosis: Pronotum with short transverse keels or fields of coarse chitin grains;
femora thickened posteriorly, apically acutely spined.
Definition: Monophyletic clade based on 12S RNA + 16S rRNA + COI + 18S rRNA
+ 28S rRNA genes phylogeny. The least inclusive clade containing Neoclytus acuminatus
(Fabricius, 1775), but not including Clytus arietis (Linnaeus, 1758).
Subordinated taxa:
1. Genus Neoclytus Thomson, 1860: 67 [67] (type species Callidium erythrocephalum
Fabricius, 1787: 156 [16] (= Callidium acuminatum Fabricius, 1775: 194 [15]);
– Neoclytus leucozonus (Laporte de Castelnau & Gory, 1836)
– Neoclytus jouteli Davis, 1904
– Neoclytus acuminatus (Fabricius, 1775)
2. Genus Zajciwiclytus, gen. nov. (type species Callidium mucronatum Fabricius,
1775: 193 [15]) – body elongated, subcylindrical, with a hair pattern of spots and transverse
stripes. Pronotum subspherical, oblong, with a wide central longitudinal band of very coarse
chitin grains. Elytra attenuated posteriorly, apically truncated, acutely spined on the outer
edge of apex. Legs very elongated. Femora gradually thickened posteriorly, apically acutely
spined. Diagnosis: Pronotum with a wide central longitudinal band of very coarse chitin
grains; elytral apex acutely spined on the outer edge. Etymology: named in honor of Dmytro
Zajciw an Ukrainian and Brazilian entomologist.
– Zajciwiclytus mucronatus (Fabricius, 1775), comb. nov.
3. Genus Cotyclytus Martins & Galileo, 2011: 127 [43] (type species Clytus curvatus
Germar, 1821: 171 [20]);
– Cotyclytus curvatus (Germar, 1821)

2.2. Subtribe Clytina, subtrib. nov.
Type genus: Clytus Laicharting, 1784: 88 [35]
Description: Body elongated, subcylindrical, with a hair pattern of spots and
transverse stripes. Head short, forehead wide, with longitudinal carinae or sulcus. Antennae
short, gradually thickened posteriorly. Pronotum subspherical, slightly oblong, uniformly
sculptured. Elytra parallel, apically truncated. Legs weak elongated. Femora gradually
thickened posteriorly.
Diagnosis: Forehead with longitudinal keels or groove; pronotum subspherical and
uniformly sculptured.
Definition: Monophyletic clade based on 12S RNA + 16S rRNA + COI + 18S rRNA
+ 28S rRNA genes phylogeny. The least inclusive clade containing Clytus arietis (Linnaeus,
1758), but not including Neoclytus acuminatus (Fabricius, 1775).
Subordinated taxa:
1. Genus Brachyclytus Kraatz, 1879: 107 [33] (type species Brachyclytus singularis
Kraatz, 1879: 107 [33]);
– Brachyclytus singularis Kraatz, 1879
2. Genus Xyloclytus Reitter, 1912: 46 [60], gen. stat. nov. (type species Clytus
chinensis Chevrolat, 1852: 416 [8])