Page 208 - Proceedings of the State Natural History Museum. Issue 37 (Lviv, 2021)

Іs clytini monophyletic? The evidence from five-gene phylogenetic … 207

Definition: Monophyletic clade based on 12S RNA + 16S rRNA + COI + 18S rRNA
+ 28S rRNA genes phylogeny. The least inclusive clade containing Chlorophorus annularis
(Fabricius, 1787), but not including Plagionotus detritus (Linnaeus, 1758).
Subordinated taxa:
1. Epiclytus Gressitt, 1935: 173 [26] (Clytus yokoyamai Kano, 1933: 275 [30])
– Epiclytus yokoyamai (Kano, 1933)
2. Genus (group I) Rhaphuma Pascoe, 1858: 240 [55], (type species Clytus
quadricolor Castelnau & Gory, 1841: 104 [36])
– Rhaphuma elongata Gressitt, 1940
3. Genus Chlorophorus Chevrolat, 1863: 290 [10], sensu nov. (type species Callidium
annulare Fabricius, 1787: 156 [16]) – Body slightly elongated, subcylindrical, with a hair
pattern of spots and transverse stripes. Head slightly elongated, clypeus well developed.
Forehead trapezoidal, widened anteriorly, with longitude medial line. Antennal bases closely
spaced. Antennae elongated, reach the second third of elytra. Typical antennal formula
rd
rd
th
th
st
1>3>5=4: 1 antennomere longer than 3 ; 3 antennomere longer than 5 ; 5 antennomere
th
equal 4 . Pronotum subspherical, slightly oblong, convex. Prosternal process narrow,
approximately 2 times wider than the edging of the procoxal cavities. Legs elongated. Outer
side of middle (and often hind) femur with a very narrow longitudinal keel. First
th
metatarsomere significantly longer then 2-5 together. Diagnosis: Forehead trapezoidal,
widened anteriorly; antennal formula 1>3>5=4; prosternal process 2 times wider than the
edging of the procoxal cavities; outer side of middle femur with a very narrow longitudinal
keel.
– Chlorophorus annularis (Fabricius, 1787)
– Chlorophorus varius (Müller, 1766)
– Chlorophorus anticemaculatus Schwarzer, 1925
– Chlorophorus annulatus (Hope, 1831)
4. Genus Clytocera Gahan, 1906: 279 (type species Clytocera chionospila Gahan,
1906: 280)
– Clytocera taiwanensis Hayashi, 1974
5. Genus Humeromaculatus Özdikmen, 2011: 537 [53], gen. stat. nov. (type species
Cerambyx figuratus Scopoli, 1763: 55 [63]) – Body elongated, subcylindrical, with a hair
pattern of spots and transverse stripes. Head elongated; clypeus well developed. Forehead
trapezoidal, widened anteriorly. Antennal bases closely spaced. Antennae elongated, reach
rd
st
the second third of elytra. Typical antennal formula 1=3=5>4: 1 antennomere equal 3 and
th
th
5 ; 5 antennomere longer than 4 . Pronotum subspherical, oblong, convex. Prosternal
th
process narrow, as wide (or narrower) as edging of the procoxal cavities. Legs elongated.
th
First metatarsomere significantly longer then 2-5 together. Diagnosis: Forehead
trapezoidal, widened anteriorly; antennal formula 1=3=5>4; pronotum oblong; prosternal
process narrow, as wide as edging of the procoxal cavities.
5.1. Subgenus Viridiphorus, subgen. nov. (type species Callidium herbstii Brahm,
1790: 148 [5]) – Body notably elongated, subcylindrical, with contrasting hair pattern of
spots. Head elongated; clypeus well developed. Forehead trapezoidal, widened anteriorly,
laterally edged with thin ribs. Antennal bases closely spaced. Antennae elongated, reach the
st
rd
second third of elytra. Typical antennal formula 1<3>5=4: 3 antennomere longer than 1
th
th
th
and 5 ; 5 antennomere equal 4 . Pronotum subspherical, notably oblong, convex. Prosternal
process narrow, as wide (or narrower) as edging of the procoxal cavities. Legs elongated.