Page 210 - Proceedings of the State Natural History Museum. Issue 37 (Lviv, 2021)
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Іs clytini monophyletic? The evidence from five-gene phylogenetic … 209
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prosternal process twice wider than edging of the procoxal cavities; 1 metatarsomere as long
th
as 2-5 together. Ethymology: σπάργανα – swaddling clothes + φέρω – to carry.
– Sparganophorus diadema (Motschulsky, 1854), comb. nov.
11. Genus (group II) Demonax Thomson, 1861: 226 [68] (type species Demonax
nigrofasciatus Thomson, 1861: 227 [68])
– Demonax transilis Bates, 1884
12. Genus Rhabdoclytus Ganglbauer, 1887 [42] (type species Clytus acutivittis
Kraatz, 1879: 111 [33])
– Rhabdoclytus acutivittis (Kraatz, 1879)
13. Genus (group III) Demonax Thomson, 1861: 226 [68] (type species Demonax
nigrofasciatus Thomson, 1861: 227 [68])
– Demonax substitutus Gressitt, 1951
– Demonax bidenticornis Hayashi, 1974
14. Genus Perderomaculatus Özdikmen, 2011: 537 [53], gen. stat. nov. (type species
Cerambyx sartor Müller, 1766, 188 [45]) – Body elongated, subcylindrical, with contrasting
hair pattern of transverse bands. Head shortened; clypeus very reduced. Forehead square.
rd
Antennal bases widely spaced. Typical antennal formula 3>5>4>1: 3 antennomere longer
th
th
th
th
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than 5 ; 5 antennomere longer than 4 ; 4 antennomere longer than 1 . The second
antennomere twice long as wide. Pronotum subspherical, slightly oblong, convex. Prosternal
process narrow, as wide as edging of the procoxal cavities. Legs elongated. First
metatarsomere longer than 2-5 together. Diagnosis: Clypeus reduced; forehead square;
th
antennal bases widely spaced; antennal formula 3>5>4>1; second antennomere twice long
as wide.
– Perderomaculatus sartor (Müller, 1766), comb. nov.
15. Genus (group III) Rhaphuma Pascoe, 1858: 240 [55] (type species Clytus
quadricolor Castelnau & Gory, 1841: 104 [36])
– Rhaphuma gracilipes (Faldermann, 1835)
Conclusions
In summary, phylogeny based on 12S rRNA 16S rRNA COI 18S rRNA 28S rRNA genes
revealed nonmonophyly of Clytini s.l., which consists two large clades: Clytini, trib. sensu
nov. and Chlorophorini, trib. nov. Both tribes are related to Anaglyptini, forming a
monophyletic supertribe Chlorophoritae, supertrib. nov. Internal phylogeny of Clytini, trib.
sensu nov. and Chlorophorini, trib. nov. remains intricate and unclear in many ways. The
current results make only a partial contribution to its understanding. While the phylogeny of
Neoclytus, Xylotrechus, Clytus, Plagionotus, Meacyllene is more or less clear, then for
Chlorophorus, Demonax, Rhaphuma it remains mostly unresolved.
1. Alonso-Zarazaga, M.A. Echinocerus Mulsant, 1862 is a valid genus (Coleoptera, Cerambycidae).
// Boletin de la Sociedad Entomológica Aragonesa, 2007 – 37 – 308-309.
2. Anisimova, M. & Gascuel, O. Approximate likelihood ratio test for branches: A fast, accurate
and powerful alternative. // Systematic Biology, 2006 – 55 (4) – 539-552.
https://doi.org/10.1080/10635150600755453
3. Bates H.W. On the Longicorn Coleoptera of Japan. // The Annals and Magazine of Natural
History, London, 1873 – 12 (69) – 193-201.