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Іs clytini monophyletic? The evidence from five-gene phylogenetic …   209

                                                                   st
            prosternal process twice wider than edging of the procoxal cavities; 1  metatarsomere as long
                 th
            as 2-5  together. Ethymology: σπάργανα – swaddling clothes + φέρω – to carry.
                  – Sparganophorus diadema (Motschulsky, 1854), comb. nov.
                  11.  Genus  (group  II)  Demonax  Thomson,  1861: 226 [68]  (type  species  Demonax
            nigrofasciatus Thomson, 1861: 227 [68])
                  – Demonax transilis Bates, 1884
                  12.  Genus  Rhabdoclytus  Ganglbauer,  1887  [42]  (type  species  Clytus  acutivittis
            Kraatz, 1879: 111 [33])
                  – Rhabdoclytus acutivittis (Kraatz, 1879)
                  13. Genus (group III) Demonax Thomson, 1861: 226 [68] (type species Demonax
            nigrofasciatus Thomson, 1861: 227 [68])
                  – Demonax substitutus Gressitt, 1951
                  – Demonax bidenticornis Hayashi, 1974
                  14. Genus Perderomaculatus Özdikmen, 2011: 537 [53], gen. stat. nov. (type species
            Cerambyx sartor Müller, 1766, 188 [45]) – Body elongated, subcylindrical, with contrasting
            hair pattern of transverse bands. Head shortened; clypeus very reduced. Forehead square.
                                                                    rd
            Antennal bases widely spaced. Typical antennal formula 3>5>4>1: 3  antennomere longer
                  th
                     th
                                              th
                                                 th
                                                                          st
            than  5 ;  5   antennomere  longer  than  4 ;  4   antennomere  longer  than  1 .  The  second
            antennomere twice long as wide. Pronotum subspherical, slightly oblong, convex. Prosternal
            process  narrow,  as  wide  as  edging  of  the  procoxal  cavities.  Legs  elongated.  First
            metatarsomere  longer  than  2-5   together.  Diagnosis:  Clypeus  reduced;  forehead  square;
                                      th
            antennal bases widely spaced; antennal formula 3>5>4>1; second antennomere twice long
            as wide.
                  – Perderomaculatus sartor (Müller, 1766), comb. nov.
                  15.  Genus  (group  III)  Rhaphuma  Pascoe,  1858:  240  [55]  (type  species  Clytus
            quadricolor Castelnau & Gory, 1841: 104 [36])
                  – Rhaphuma gracilipes (Faldermann, 1835)

               Conclusions

               In summary, phylogeny based on 12S rRNA 16S rRNA COI 18S rRNA 28S rRNA genes
            revealed nonmonophyly of Clytini s.l., which consists two large clades: Clytini, trib. sensu
            nov.  and  Chlorophorini,  trib.  nov.  Both  tribes  are  related  to  Anaglyptini,  forming  a
            monophyletic supertribe Chlorophoritae, supertrib. nov. Internal phylogeny of Clytini, trib.
            sensu nov. and Chlorophorini, trib. nov. remains intricate and unclear in many ways. The
            current results make only a partial contribution to its understanding. While the phylogeny of
            Neoclytus,  Xylotrechus,  Clytus,  Plagionotus,  Meacyllene  is  more  or  less  clear,  then  for
            Chlorophorus, Demonax, Rhaphuma it remains mostly unresolved.

              1.  Alonso-Zarazaga, M.A. Echinocerus Mulsant, 1862 is a valid genus (Coleoptera, Cerambycidae).
                 // Boletin de la Sociedad Entomológica Aragonesa, 2007 – 37 – 308-309.
              2.  Anisimova, M. & Gascuel, O. Approximate likelihood ratio test for branches: A fast, accurate
                 and  powerful  alternative.  //  Systematic  Biology,  2006  –  55  (4)  –  539-552.
                 https://doi.org/10.1080/10635150600755453
              3.  Bates  H.W.  On  the  Longicorn Coleoptera  of  Japan.  //  The  Annals  and  Magazine  of  Natural
                 History, London, 1873 – 12 (69) – 193-201.
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